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Gin's monkey wrench-wielding Doom Monkeys, Uka Uka's spear-chucking Voodoo Bunnies, and Nina Cortex's Brat Girls. The later enemies include Tiny Tiger's Southern-accented Koo-alas, Doctor N. These characters appear in the first four episodes of the game, and are replaced by other enemies as the game progresses. The first fodders encountered are Neo Cortex's Ratnicians, which appear as bipedal, chihuahua-like animals dressed as laboratory researchers. In Crash: Mind Over Mutant, they are stationed as guards in the Space Head. Sanity Island as wild titans, they soon became Dr. Each fodder is affiliated with a certain villain in the game. Aku Aku, Crash of the Titans The Stench is a titan hybrid of a vulture and a skunk that has long-ranged stink bomb attacks. Aiding the villains are a number of small fodder enemies that attempt to hinder Crash in his journey. Dingodile, another minion of Cortex, exclusively appears in the DS and Game Boy Advance versions of Crash of the Titans as a boss character. Gin, who is Cortex's hysterical right-hand man, and Tiny Tiger. Uka Uka, who is also absent in the Nintendo DS version, is behind Cortex's plot for world domination after tiring of Cortex's incompetence, Uka replaces him with his cyborg niece, Nina Cortex. Charactersĭoctor Neo Cortex is a mad scientist who captures Coco and Aku Aku and wants to use stolen Mojo to create an army of mutants. The final moments of the game take place in the antagonist's Liberace-style lair, and finally inside the Doominator. This tree holds up an entire house owned by Uka Uka, one of the major antagonists of the series. The penultimate area is a gigantic tree located in the middle of the island. Gin's missile factory, which resembles the Statue of Liberty. The next few episodes take place on an artillery-bombarded beach, where Crash infiltrates Doctor N. Later in the game, Crash travels through a wood-cutting and mining area and a land flowing with lava that features giant steel mosquitoes sucking minerals out of the ground.

The story starts in the jungle near Crash's residence and leads to an ancient temple. The game moves through five major themes taking place on Crash's homeland, the Wumpa Islands. Like the first Crash Bandicoot game, episodes must be played in order, although the player can replay any episode that has been completed. Each level is referred to as an 'episode', and each has a distinctive subtitle that is usually a punning reference to popular culture (such as 'The Blizzard of Claws'). The setting of Crash of the Titans features twenty levels.